{"id":685,"date":"2015-03-18T00:06:15","date_gmt":"2015-03-17T15:06:15","guid":{"rendered":"http:\/\/www.nibb.ac.jp\/plantdic\/blog\/?p=685"},"modified":"2015-03-18T23:16:42","modified_gmt":"2015-03-18T14:16:42","slug":"rotated-or-resupinated-leaves-%e8%91%89%e3%81%ae%e3%81%ad%e3%81%98%e3%82%8c%e3%82%84%e5%8f%8d%e8%bb%a2","status":"publish","type":"post","link":"https:\/\/www.nibb.ac.jp\/plantdic\/blog\/?p=685","title":{"rendered":"rotated or resupinated leaves \u8449\u306e\u306d\u3058\u308c\u3084\u53cd\u8ee2"},"content":{"rendered":"

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Albuca namaquensis<\/em>\u306e\u8449\u306f\u57fa\u90e8\u5074\u304b\u3089\u5148\u7aef\u5074\u306b\u898b\u308b\u3068\u53cd\u6642\u8a08\u56de\u308a\u306b\u3088\u3058\u308c\u3066\u3044\u308b\u3002\u3053\u306e\u3088\u3058\u308c\u306f\u3001\u8449\u306e\u5de6\u53f3\u306e\u6210\u9577\u306e\u5ea6\u5408\u3044\u304c\u7570\u306a\u308b\u305f\u3081\u3060\u3068\u8003\u3048\u3089\u308c\u308b\u3002\u3069\u3093\u306a\u690d\u7269\u3082\u8449\u306f\u5de6\u53f3\u975e\u5bfe\u79f0\u3067\u3042\u308b\u3002\u8449\u306e\u975e\u5bfe\u79f0\u6027\u306b\u306f\uff12\u30bf\u30a4\u30d7\u306e\u6a5f\u69cb\u304c\u3042\u308b\uff08\u7dcf\u8aac\u3068\u3057\u3066 Chitwood et al. 2012)\u3002\u4e00\u3064\u306f\u8449\u5e8f\u306b\u4f9d\u5b58\u3057\u305f\u975e\u5bfe\u79f0\u6027\u3067\u3042\u308b\u3002\u3053\u308c\u306f\u3001\u8449\u539f\u57fa\u5185\u3067\u3001\u30aa\u30fc\u30ad\u30b7\u30f3\u6fc3\u5ea6\u304c\u524d\u306e\u8449\u306e\u5074\u304c\u6b21\u306e\u8449\u306e\u5074\u3088\u308a\u3082\u9ad8\u304f\u3001\u305d\u308c\u3086\u3048\u3001\u524d\u306e\u8449\u5074\u306e\u8449\u8eab\u304c\u9006\u5074\u3088\u308a\u3082\u5927\u304d\u304f\u306a\u308b\u3002\u3082\u3046\u4e00\u3064\u306e\u30bf\u30a4\u30d7\u306f\u3001\u8449\u5e8f\u306b\u4f9d\u5b58\u3057\u306a\u3044\u975e\u5bfe\u79f0\u6027\u3067\u3001\u907a\u4f1d\u7684\u306b\u6c7a\u307e\u3063\u3066\u3044\u308b\u3082\u306e\u3067\u3042\u308b\u3002\u4f8b\u3048\u3070\u3001\u30a2\u30ea\u30b9\u30c8\u30ed\u30e1\u30ea\u30a2\u3084\u30b9\u30a4\u30bb\u30f3\u306e\u8449\u306f\u3001\u5411\u304b\u3063\u3066\u53f3\u5074\u304c\u826f\u304f\u80b2\u3061\u3001\u7d50\u679c\u3068\u3057\u3066\u53cd\u6642\u8a08\u56de\u308a\u306b\u56de\u8ee2\u3057\u305f\u8449\u304c\u3067\u304d\u308b\u3002Albuca namaquensis<\/em>\u306e\u5834\u5408\u3082\u3001\u3069\u306e\u500b\u4f53\u3082\u53cd\u6642\u8a08\u56de\u308a\u306b\u3088\u3058\u308c\u3066\u3044\u308b\u306e\u3067\u3001\u8449\u5e8f\u306b\u4f9d\u5b58\u3057\u306a\u3044\u30bf\u30a4\u30d7\u3067\u3042\u308b\u3002\u3069\u306e\u3088\u3046\u306b\u5206\u5b50\u6a5f\u69cb\u3067\u5de6\u53f3\u975e\u5bfe\u79f0\u6027\u3092\u7523\u307f\u51fa\u3057\u3001\u56de\u65cb\u3057\u305f\u8449\u3092\u5f62\u6210\u3059\u308b\u306e\u304b\u306f\u308f\u304b\u3063\u3066\u3044\u306a\u3044\u3002<\/p>\n

There are two types of mechanisms to cause asymmetry in leaves (summarized in Chitwood et al. 2012). One is phyllotaxis-dependent asymmetry, which is caused by asymmetric distribution of auxin in a leaf primordium. Auxin is more accumulated in a previous leaf primordium side than in a next coming leaf primordium, which results in a larger leaf blade in the former side than the latter side. The other is phyllotaxis\u2013independent asymmetry, which is usually inherent. In Alstroemeria and Narcissus, leaves are resupinate or rotate in a counter-clockwise direction. Leaves of Albuca namaquensis<\/em> rotates also in a counter-clockwise direction. The molecular mechanisms of phyllotaxis-independent asymmetry is unknown.<\/p>\n

Chitwood, D.H. Naylor, D.t., Thammapichai, P., Weeger, A.C.S., Headland, L.R., and Sinha, N.R. 2012. Conflict between intrinsic leaf asymmetry and phyllotaxis in the resupinate leaves of Alstroemeria psittacina. Front. Plant Sci. 3: 1-11.<\/p>\n

\"\"<\/a><\/p>\n

\u8449\u306e\u306d\u3058\u308c\u306f\u5358\u5b50\u8449\u985e\u306e\u30a2\u30e4\u30e1\u79d1\uff08\u30ad\u30b8\u30ab\u30af\u30b7\u76ee\uff09\u3001\u30d2\u30ac\u30f3\u30d0\u30ca\u79d1\uff08\u30ad\u30b8\u30ab\u30af\u30b7\u76ee\uff09\u3001\u30eb\u30ba\u30ea\u30a2\u30ac\u79d1\uff08\u30e6\u30ea\u76ee\uff09\u3001\u30a2\u30ea\u30b9\u30c8\u30ed\u30e1\u30ea\u30a2\u79d1\uff08\u30e6\u30ea\u76ee\uff09\u3001\u30a4\u30cd\u79d1\uff08\u30a4\u30cd\u76ee\uff09\u3067\u898b\u3089\u308c\u308b\u3002\u4e0d\u601d\u8b70\u306a\u3053\u3068\u306b\u3001\u3069\u306e\u5834\u5408\u3082\u8449\u306f\u53cd\u6642\u8a08\u56de\u308a\u306b\u306d\u3058\u308c\u308b\u3002\u30e6\u30fc\u30ab\u30ea\u306a\u3069\u306e\u53cc\u5b50\u8449\u985e\u3082\u3084\u306f\u308a\u53cd\u6642\u8a08\u56de\u308a\u306b\u306d\u3058\u308c\u308b\u3002\u3068\u3053\u308d\u304c\u3001\u30b3\u30d0\u30f3\u30bd\u30a6\u306f\u4f8b\u5916\u7684\u306b\u53f3\u306b\u306d\u3058\u308c\u308b\uff08\u718a\u6fa4 1979)\u3002\u7406\u7531\u306f\u4e0d\u660e\u3067\u3042\u308b\u3002<\/p>\n

\u30eb\u30ba\u30ea\u30a2\u30ac\u79d1\u3084\u30a2\u30ea\u30b9\u30c8\u30ed\u30e1\u30ea\u30a2\u79d1\u306e\u7a2e\u3067\u306f\u3001\u8449\u304c\u306d\u3058\u308c\u304c\u8457\u3057\u304f\u3001\u53cd\u8ee2\u3057\u3066\u3057\u307e\u3046\u3002\u3057\u304b\u3082\u3001\u6c17\u5b54\u306e\u5206\u5e03\u3001\u8449\u306e\u8272\u3001\u30af\u30c1\u30af\u30e9\u306e\u91cf\u3001\u67f5\u72b6\u7d44\u7e54\u3068\u6d77\u7dbf\u72b6\u7d44\u7e54\u306e\u5206\u5e03\u3082\u901a\u5e38\u306e\u8449\u3068\u8f03\u3079\u3066\u9006\u8ee2\u3059\u308b\uff08\u718a\u6fa4 1979)\u3002\u3069\u3046\u3057\u3066\u9006\u8ee2\u3057\u3066\u3057\u307e\u3046\u306e\u304b\u3001\u305d\u3057\u3066\u3001\u53cd\u6642\u8a08\u56de\u308a\u306a\u306e\u304b\u3001\u7406\u7531\u306f\u4e0d\u660e\u3067\u3042\u308b\u3002<\/p>\n

Leaves are rotated in some species of different monocot families including Iridaceae (Asparagales), Amaryllidaceae (Asparagales), Luzuriagaceae (Liliales), Alstroemeriaceae (Liliales), and Poaceae (Poales). Curiously, leaves always rotate in a counter-clockwise direction except few exceptions including Briza maxima. The reason is unknown.<\/p>\n

In species in the Luzuriagaceae and Alstroemeriaceae, Hakonechloa macra, Elymus spp., and so on, leaves are resupinated and characters of leaves including stomata distribution, cuticle deposition, coloration, organization of palisade and spongy cells are reversed between adaxial and abaxial sides.<\/p>\n

\u718a\u6fa4\u6b63\u592b (Kumazawa, M.) 1979 \u690d\u7269\u5668\u5b98\u5b66 (Plant Organography) \u88f3\u83ef\u623f (Publisher: Shoka-Bo)<\/p>\n

\"\"<\/a><\/p>\n

\u30a4\u30cd\u79d1\u306e\u30a6\u30e9\u30cf\u30b0\u30b5\u5c5e\u3001\u30ab\u30e2\u30b8\u30b0\u30b5\u5c5e\u3001\u30ab\u30bc\u30af\u30b5\u5c5e\u306e\u8449\u306f\u534a\u6642\u8a08\u5468\u308a\u306b\u8868\u88cf\u304c\u9006\u8ee2\u3057\u3066\u3044\u308b\u3002\u30b3\u30d0\u30f3\u30bd\u30a6\u306f\u4f8b\u5916\u3067\u3001\u6642\u8a08\u56de\u308a\u306b\u9006\u8ee2\u3057\u3066\u3044\u308b\uff08\u718a\u6fa4 1979)\u3002<\/p>\n

Leaves of some Poaceae genera including Hakonechloa macra<\/em>, Elymus<\/em> spp., Eragrostis<\/em> spp. and so on are resupinated in a counter-clockwise direction. Briza maxima<\/em> with resupinated leaves with a clockwise direction is exceptional.<\/p>\n

\"\"<\/a><\/p>\n

\"\"<\/a><\/p>\n

Gethyllis<\/em>\u5c5e\u306f\u30a2\u30d5\u30ea\u30ab\u5357\u90e8\u306b\u7d0432\u7a2e\u304c\u5206\u5e03\u3059\u308b\uff08Meerow and Snijman 1998)\u3002\u8449\u306f\u53cd\u6642\u8a08\u56de\u308a\u306b\u56de\u65cb\u3059\u308b\u3002<\/p>\n

Gethyllis<\/em> is composed of approximately 32 species distributed in Southern Africa (Meerow and Snijman 1998). Leaves rotate in a counter-clockwise direction.<\/p>\n

Meerow, A.W. and Snijman, D.A. 1998. Amaryllidaceae. In Kubitzki, K. The Families and Genera of Vascular Plants. III Flowering Plants Monocotyledons Lilianae (except Orchidaceae). Pp. 83-110<\/p>\n

 <\/p>\n","protected":false},"excerpt":{"rendered":"

Albuca namaquensis\u306e\u8449\u306f\u57fa\u90e8\u5074\u304b\u3089\u5148\u7aef\u5074\u306b\u898b\u308b\u3068\u53cd\u6642\u8a08\u56de\u308a\u306b\u3088 … \u7d9a\u304d\u3092\u8aad\u3080 →<\/span><\/a><\/p>\n","protected":false},"author":1,"featured_media":0,"comment_status":"closed","ping_status":"closed","sticky":false,"template":"","format":"standard","meta":[],"categories":[146],"tags":[],"_links":{"self":[{"href":"https:\/\/www.nibb.ac.jp\/plantdic\/blog\/index.php?rest_route=\/wp\/v2\/posts\/685"}],"collection":[{"href":"https:\/\/www.nibb.ac.jp\/plantdic\/blog\/index.php?rest_route=\/wp\/v2\/posts"}],"about":[{"href":"https:\/\/www.nibb.ac.jp\/plantdic\/blog\/index.php?rest_route=\/wp\/v2\/types\/post"}],"author":[{"embeddable":true,"href":"https:\/\/www.nibb.ac.jp\/plantdic\/blog\/index.php?rest_route=\/wp\/v2\/users\/1"}],"replies":[{"embeddable":true,"href":"https:\/\/www.nibb.ac.jp\/plantdic\/blog\/index.php?rest_route=%2Fwp%2Fv2%2Fcomments&post=685"}],"version-history":[{"count":3,"href":"https:\/\/www.nibb.ac.jp\/plantdic\/blog\/index.php?rest_route=\/wp\/v2\/posts\/685\/revisions"}],"predecessor-version":[{"id":697,"href":"https:\/\/www.nibb.ac.jp\/plantdic\/blog\/index.php?rest_route=\/wp\/v2\/posts\/685\/revisions\/697"}],"wp:attachment":[{"href":"https:\/\/www.nibb.ac.jp\/plantdic\/blog\/index.php?rest_route=%2Fwp%2Fv2%2Fmedia&parent=685"}],"wp:term":[{"taxonomy":"category","embeddable":true,"href":"https:\/\/www.nibb.ac.jp\/plantdic\/blog\/index.php?rest_route=%2Fwp%2Fv2%2Fcategories&post=685"},{"taxonomy":"post_tag","embeddable":true,"href":"https:\/\/www.nibb.ac.jp\/plantdic\/blog\/index.php?rest_route=%2Fwp%2Fv2%2Ftags&post=685"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}