{"id":1577,"date":"2026-06-15T13:20:21","date_gmt":"2026-06-15T04:20:21","guid":{"rendered":"https:\/\/www.nibb.ac.jp\/plantdic\/blog\/?p=1577"},"modified":"2026-06-15T13:20:21","modified_gmt":"2026-06-15T04:20:21","slug":"%e3%82%b0%e3%83%ad%e3%83%aa%e3%82%aa%e3%82%b5%e5%b1%9e-gloriosa","status":"publish","type":"post","link":"http:\/\/www.nibb.ac.jp\/plantdic\/blog\/?p=1577","title":{"rendered":"\u30b0\u30ed\u30ea\u30aa\u30b5\u5c5e Gloriosa"},"content":{"rendered":"\n
\"\"<\/a>
\u30b0\u30ed\u30ea\u30aa\u30b5\u5c5e<\/strong>
Gloriosa<\/em><\/strong>
Gloriosa superba<\/em>: Cropped from a photograph by Chayant Gonsalves, https:\/\/www.inaturalist.org\/photos\/634639040, CC BY-NC
Source: POWO (2026). Plants of the World Online<\/em>. Facilitated by the Royal Botanic Gardens, Kew. Published on the Internet; https:\/\/powo.science.kew.org\/<\/a>. Retrieved 14 June 2026. Distribution data from Kew Backbone Distributions (WCVP). \u00a9 World Checklist of Vascular Plants, licensed under CC BY 3.0.<\/figcaption><\/figure>\n\n\n\n

\u30b0\u30ed\u30ea\u30aa\u30b5\u5c5e<\/strong> Gloriosa<\/strong><\/em><\/p>\n\n\n\n

\u30b0\u30ed\u30ea\u30aa\u30b5\u5c5e\u306f\u30a4\u30cc\u30b5\u30d5\u30e9\u30f3\u79d1\u306b\u5c5e\u3057\u300111\u7a2e\u304c\u77e5\u3089\u308c\u3066\u304a\u308a\u3001\u30a2\u30d5\u30ea\u30ab\u3001\u30a2\u30b8\u30a2\u3092\u4e2d\u5fc3\u306b\u5206\u5e03\u3059\u308b\uff08POWO\uff09\u3002<\/p>\n\n\n\n

Gloriosa<\/em> belongs to Colchicaceae and comprises 11 known species, distributed mainly in Africa and Asia (POWO).<\/p>\n\n\n\n

\"\"<\/a><\/figure>\n\n\n\n
\"\"<\/a>
Gloriosa rigidifolia<\/em>: Cropped from a photo by Manuel R Popp, https:\/\/www.inaturalist.org\/photos\/174698901, CC BY
Gloriosa rigidifolia<\/em>: Cropped from a photo by Manuel R Popp, https:\/\/www.inaturalist.org\/photos\/169349621, CC BY<\/figcaption><\/figure>\n\n\n\n

Gloriosa\u306b\u306f\u82b1\u88ab\u7247\u304c\u5f37\u304f\u53cd\u308a\u8fd4\u308a\u3001\u82b1\u67f1\u304c\u307b\u307c\u76f4\u89d2\u306b\u66f2\u304c\u308b\u7a2e\uff08\u4f8b\u3048\u3070Gloriosa superba<\/em>\uff09\u3068\u3001\u82b1\u88ab\u7247\u304c\u5f37\u304f\u53cd\u308a\u8fd4\u3089\u305a\u3001\u82b1\u67f1\u304c\u307b\u3068\u3093\u3069\u66f2\u304c\u3089\u306a\u3044\u7a2e\uff08\u4f8b\u3048\u3070Gloriosa rigidifolia<\/em>\uff09\u304c\u3042\u308a\u3001\u5f8c\u8005\u306f\u304b\u3064\u3066\u30ea\u30c3\u30c8\u30cb\u30a2\u5c5eLittonia<\/em>\u3068\u3057\u3066\u6271\u308f\u308c\u305f\u3053\u3068\u3082\u3042\u3063\u305f\u304c\u3001\u5206\u5b50\u7cfb\u7d71\u89e3\u6790\u306b\u3088\u308a\u30b0\u30ed\u30ea\u30aa\u30b5\u5c5e\u3068\u5358\u7cfb\u7d71\u7fa4\u3092\u5f62\u6210\u3059\u308b\u3053\u3068\u304c\u308f\u304b\u308a\u3001\u73fe\u5728\u3067\u306f\u30b0\u30ed\u30ea\u30aa\u30b5\u5c5e\u306b\u7d71\u5408\u3055\u308c\u3066\u3044\u308b (Thi et al. 2013)\u3002<\/p>\n\n\n\n

Gloriosa<\/em> includes species with strongly reflexed tepals and a style bent almost at a right angle, such as Gloriosa superba<\/em>, as well as species with tepals that are not strongly reflexed and a style that is scarcely bent, such as Gloriosa rigidifolia<\/em>. The latter were once treated as members of the genus Littonia<\/em>, but molecular phylogenetic analyses showed that they form a monophyletic group with Gloriosa<\/em>. They are therefore now included in Gloriosa<\/em> (Thi et al. 2013).<\/p>\n\n\n\n

\"\"<\/a>
\u30b0\u30ed\u30ea\u30aa\u30b5\u306e\u307b\u307c\u76f4\u89d2\u306b\u66f2\u304c\u308b\u96cc\u305a\u3044<\/strong>
a style of Gloriosa superba<\/em> bent almost at a right angle<\/strong><\/figcaption><\/figure>\n\n\n\n

\u30b0\u30ed\u30ea\u30aa\u30b5\u306e\u307b\u307c\u76f4\u89d2\u306b\u66f2\u304c\u308b\u96cc\u305a\u3044<\/strong><\/p>\n\n\n\n

a style of <\/strong>Gloriosa superba<\/em><\/strong> bent almost at a right angle<\/strong><\/p>\n\n\n\n

Gloriosa superba<\/em>\u306e\u82b1\u67f1\u306f\u3001\u767a\u751f\u904e\u7a0b\u3067\u57fa\u90e8\u304c\u5411\u8ef8\u5074\uff08\u82b1\u306e\u4e0a\u5074\uff09\u3078\u307b\u307c\u76f4\u89d2\u306b\u5c48\u66f2\u3059\u308b\u3002\u5b50\u623f\u3067\u306f\u5411\u8ef8\u5074\u306e\u5b50\u623f\u5ba4\u306e\u5927\u304d\u3055\u304c\u80cc\u8ef8\u5074\u306e\u5b50\u623f\u5ba4\u3088\u308a\u3082\u5c0f\u3055\u304f\u3001\u857e\u306e\u6bb5\u968e\u3067\u3059\u3067\u306b\u5c48\u66f2\u304c\u8a8d\u3081\u3089\u308c\u308b\u3002\u67f1\u982d\u306f\u846f\u3088\u308a\u7a81\u51fa\u3059\u308b\u305f\u3081\u3001\u4ed6\u500b\u4f53\u306e\u82b1\u7c89\u3092\u4ed8\u7740\u3057\u305f\u9001\u7c89\u6606\u866b\u3067\u3042\u308b\u30b7\u30ed\u30c1\u30e7\u30a6\u79d1\u306e\u30c1\u30e7\u30a6\u985e\u306f\u3001\u8a2a\u82b1\u6642\u306b\u307e\u305a\u67f1\u982d\u306b\u63a5\u89e6\u3057\u3084\u3059\u3044\u3002\u305d\u306e\u7d50\u679c\u3001\u81ea\u5bb6\u53d7\u7c89\u306e\u5272\u5408\u304c\u4f4e\u4e0b\u3057\u3001\u4ed6\u5bb6\u53d7\u7c89\u306e\u5272\u5408\u304c\u9ad8\u307e\u308b\u3068\u8003\u3048\u3089\u308c\u308b\uff08Daniels et al. 2020)\u3002\u540c\u69d8\u306e\u96cc\u305a\u3044\u306e\u5c48\u66f2\u306f\u3001Lilium martagon<\/em>\u3067\u3082\u89b3\u5bdf\u3055\u308c\u3066\u3044\u308b\uff08Brantjes and Bos 1980, Corbera et al. 2018)\u3002<\/p>\n\n\n\n

In Gloriosa superba<\/em>, the style becomes bent almost at a right angle toward the adaxial side, i.e. the upper side of the flower, during development. In the ovary, the adaxial locule is smaller than the abaxial locule, and the curvature is already evident at the bud stage. Because the stigma protrudes beyond the anthers, pierid butterflies carrying pollen from other individuals are likely to contact the stigma first when visiting the flower. This is thought to reduce the frequency of self-pollination and increase the frequency of cross-pollination (Daniels et al. 2020). A similar curvature of the pistil has also been observed in Lilium martagon<\/em> (Brantjes and Bos 1980; Corbera et al. 2018).<\/p>\n\n\n\n

\"\"<\/a>
\u30b0\u30ed\u30ea\u30aa\u30b5\u306e\u9577\u3044\u53e3\u543b\u7528\u306e\u871c\u817a<\/strong>
nectaries adapted for long proboscides in Gloriosa superba<\/em><\/strong><\/figcaption><\/figure>\n\n\n\n

\u30b0\u30ed\u30ea\u30aa\u30b5\u306e\u9577\u3044\u53e3\u543b\u7528\u306e\u871c\u817a<\/strong><\/p>\n\n\n\n

nectaries adapted for long proboscides in <\/strong>Gloriosa superba<\/em><\/strong><\/p>\n\n\n\n

\u30e6\u30ea\u76ee\u306e\u5171\u6709\u6d3e\u751f\u5f62\u8cea\u306f\u3001\u82b1\u88ab\u7247\u4e0a\u306b\u82b1\u88ab\u871c\u817a\u3092\u6301\u3064\u3053\u3068\u3067\u3042\u308b\u3002Gloriosa superba<\/em>\u306e\u871c\u817a\u306f\u82b1\u88ab\u7247\u306e\u5411\u8ef8\u5074\u57fa\u90e8\u306b\u3042\u308a\u3001\u30c1\u30e7\u30a6\u985e\u306f\u5de6\u306e\u5199\u771f\u306e\u82b1\u5f01\u5411\u8ef8\u5074\u306e\u5165\u53e3\u304b\u3089\u53e3\u543b\u3092\u5dee\u3057\u8fbc\u307f\uff08\u70b9\u7dda\uff09\u3001\u82b1\u88ab\u7247\u57fa\u90e8\u306b\u3042\u308b\u871c\u817a\u304b\u3089\u5438\u871c\u3059\u308b\u3002\u871c\u817a\u3078\u306e\u5165\u53e3\u306f\u3001\u53e3\u543b\u306e\u77ed\u3044\u6606\u866b\u306e\u5438\u871c\u3092\u5236\u9650\u3057\u3001\u53e3\u543b\u306e\u9577\u3044\u30c1\u30e7\u30a6\u985e\u306b\u3088\u308b\u8a2a\u82b1\u3092\u4fc3\u9032\u3057\u3066\u3044\u308b\u3068\u8003\u3048\u3089\u308c\u308b\uff08Daniels et al. 2020)\u3002\u30c1\u30e7\u30a6\u985e\u304c\u53cd\u308a\u8fd4\u3063\u305f\u82b1\u88ab\u7247\u306e\u57fa\u90e8\u306b\u3042\u308b\u871c\u817a\u3078\u30a2\u30d7\u30ed\u30fc\u30c1\u3059\u308b\u3068\u3001\u7fc5\u306e\u4e0b\u9762\u304c\u846f\u306b\u63a5\u89e6\u3057\u3001\u82b1\u7c89\u304c\u4ed8\u7740\u3059\u308b\uff08Daniels et al. 2020)\u3002 A shared derived character of Liliales is the presence of tepal nectaries on the tepals. In Gloriosa superba<\/em>, the nectaries are located at the adaxial base of the tepals. Butterflies insert their proboscides through the entrance on the adaxial side of the tepal shown in the photograph on the left (dotted line) and feed on nectar from the nectaries at the tepal base. The entrance to the nectary is thought to restrict nectar access by insects with short proboscides and to promote visitation by long-proboscid butterflies (Daniels et al. 2020). When butterflies approach the nectaries at the base of the reflexed tepals, the undersides of their wings come into contact with the anthers and become dusted with pollen (Daniels et al. 2020).<\/p>\n\n\n\n

\"\"<\/a>
\u30b0\u30ed\u30ea\u30aa\u30b5\u306e\u82b1\u88ab\u306e\u8272\u5f69<\/strong>
Perianth coloration in Gloriosa<\/em><\/strong><\/figcaption><\/figure>\n\n\n\n

\u30b0\u30ed\u30ea\u30aa\u30b5\u306e\u82b1\u88ab\u306e\u8272\u5f69<\/strong><\/p>\n\n\n\n

Perianth coloration in <\/strong>Gloriosa<\/em><\/strong><\/p>\n\n\n\n

\u30b0\u30ed\u30ea\u30aa\u30b5\u306e\u82b1\u88ab\u7247\u306b\u898b\u3089\u308c\u308b\u30aa\u30ec\u30f3\u30b8\u8272\u304b\u3089\u8d64\u8272\u3068\u9ec4\u8272\u306e\u6a21\u69d8\u306f\u3001\u30c1\u30e7\u30a6\u985e\u306b\u8a8d\u8b58\u3055\u308c\u3084\u3059\u304f\uff08Weiss 1997)\u3001V\u5b57\u578b\u306e\u6a21\u69d8\u306f\u871c\u817a\u3078\u306e\u8a98\u5f15\u8def\u3068\u3057\u3066\u6a5f\u80fd\u3057\u3066\u3044\u308b\u53ef\u80fd\u6027\u304c\u3042\u308b\uff08Daniels et al. 2020)\u3002 The orange-to-red and yellow pattern seen on the tepals of Gloriosa<\/em> is easily recognized by butterflies (Weiss 1997), and the V-shaped pattern may function as a guide leading them to the nectaries (Daniels et al. 2020).<\/p>\n\n\n\n

\"\"<\/a>
\u30b0\u30ed\u30ea\u30aa\u30b5\u306e\u8449\u5148\u306e\u5dfb\u304d\u3072\u3052<\/strong>
Leaf-tip tendrils of Gloriosa superba<\/em><\/strong><\/figcaption><\/figure>\n\n\n\n

\u30b0\u30ed\u30ea\u30aa\u30b5\u306e\u8449\u5148\u306e\u5dfb\u304d\u3072\u3052<\/strong><\/p>\n\n\n\n

Leaf-tip tendrils of <\/strong>Gloriosa superba<\/em><\/strong><\/p>\n\n\n\n

\u30b0\u30ed\u30ea\u30aa\u30b5\u306e\u8449\u8eab\u90e8\u306e\u5148\u7aef\u5074 (\u5199\u771f\u306e\u9ed2\u8272\u77e2\u5370) \u3067\u306f\u8868\u88cf\u306e\u533a\u5225\u304c\u660e\u77ad\u3067\u3042\u308b\u3002\u4e00\u65b9\u3001\u5dfb\u304d\u3072\u3052\u3068\u306a\u308b\u8449\u5148\u90e8\uff08\u8d64\u8272\u77e2\u5370\uff09\u306f\u3001\u8449\u8eab\u90e8\u3068\u7570\u306a\u308a\u3001\u3084\u3084\u592a\u304f\u7d30\u9577\u3044\u5186\u67f1\u72b6\u3068\u306a\u308b\u3002\u3053\u306e\u305f\u3081\u5916\u5f62\u306f\u653e\u5c04\u76f8\u79f0\u7684\u306b\u898b\u3048\u308b\u304c\u3001\u7dad\u7ba1\u675f\u306f\u80cc\u8179\u6027\u3092\u4fdd\u6301\u3057\u3066\u3044\u308b (Arber 1920)\u3002\u5dfb\u304d\u3072\u3052\u306e\u5148\u7aef\u306f\u786c\u304f\u68d8\u72b6\u3068\u306a\u308a\u3001\u4ed6\u7269\u306b\u5f15\u3063\u304b\u304b\u308a\u3084\u3059\u3044\u3002\u8449\u5148\u306f\u3001\u4ed6\u7269\u306b\u63a5\u89e6\u3059\u308b\u304b\u3069\u3046\u304b\u306b\u304b\u304b\u308f\u3089\u305a\u3001\u4f38\u9577\u3092\u7d9a\u3051\u308b\u3002 In the distal part of the leaf blade of Gloriosasuperba <\/em>(black arrow in the photograph), the distinction between the adaxial and abaxial surfaces is clear. In contrast, the leaf-tip region that develops into a tendril (red arrow) differs from the leaf blade in becoming a somewhat thickened, slender cylindrical structure. Thus, its external morphology appears radially symmetrical, but the vascular bundles retain dorsiventrality (Arber 1920). The tip of the tendril becomes hard and spine-like, making it easy to catch on surrounding objects. The leaf tip continues to elongate regardless of whether it comes into contact with surrounding objects.<\/p>\n\n\n\n

\"\"<\/a>
\u30b0\u30ed\u30ea\u30aa\u30b5\u5c5e\u306e\u7403\u830e<\/strong>
Corms of Gloriosa<\/em><\/strong><\/figcaption><\/figure>\n\n\n\n

\u30b0\u30ed\u30ea\u30aa\u30b5\u306e\u7403\u830e<\/strong><\/p>\n\n\n\n

Corms of <\/strong>Gloriosa superba<\/em><\/strong><\/p>\n\n\n\n

\u30a4\u30cc\u30b5\u30d5\u30e9\u30f3\u79d1\u306e\u690d\u7269\u306f\u3001\u7403\u830e\u307e\u305f\u306f\u5730\u4e0b\u830e\u3092\u5f62\u6210\u3059\u308b(Nordenstam 1998)\u3002\u5f53\u5e74\u30b7\u30e5\u30fc\u30c8\u306e\u6700\u521d\u306e2\u8449\u306e\u814b\u82bd\u306f\u3001\u7fcc\u5e74\u306e\u30b7\u30e5\u30fc\u30c8\u306e\u830e\u9802\u3068\u306a\u308b(Nordenstam 1998, \u3002\u5f53\u5e74\u30b7\u30e5\u30fc\u30c8\u81ea\u4f53\u3082\u814b\u82bd\u306b\u7531\u6765\u3059\u308b\u305f\u3081\u3001\u7b2c1\u8449\u304a\u3088\u3073\u7b2c2\u8449\u306f\u524d\u8449\u306b\u76f8\u5f53\u3057\u3001\u3053\u308c\u3089\u524d\u8449\u306e\u814b\u82bd\u304c\u7fcc\u5e74\u306e\u30b7\u30e5\u30fc\u30c8\u306e\u830e\u9802\u306b\u306a\u308b\u3002\u4e00\u65b9\u3001\u524d\u8449\u4ee5\u5916\u306e\u8449\u306e\u814b\u82bd\u306f\u3001\u5730\u4e0a\u90e8\u3067\u5206\u679d\u3092\u5f62\u6210\u3059\u308b\u3002\u7b2c1\u8449\u306e\u814b\u82bd\u306f\u66f4\u65b0\u82bdinnovation bud\u3068\u306a\u308a\u3001\u7fcc\u5e74\u306b\u4f38\u9577\u3059\u308b\u3002\u7b2c2\u8449\u306e\u814b\u82bd\u306f\u4e88\u5099\u82bdreserve bud\u3068\u306a\u308a\u3001\u901a\u5e38\u306f\u66f4\u65b0\u82bd\u304c\u640d\u50b7\u307e\u305f\u306f\u67af\u6b7b\u3057\u305f\u5834\u5408\u306b\u4f38\u9577\u3059\u308b\u3002\u305f\u3060\u3057\u3001\u74b0\u5883\u6761\u4ef6\u3084\u7a2e\u306b\u3088\u3063\u3066\u306f\u3001\u66f4\u65b0\u82bd\u3068\u4e88\u5099\u82bd\u306e\u4e21\u65b9\u304c\u4f38\u9577\u3059\u308b\u3053\u3068\u3082\u3042\u308b\u3002\u30a4\u30cc\u30b5\u30d5\u30e9\u30f3\u3068\u30b0\u30ed\u30ea\u30aa\u30b5\u306e\u7403\u830e\u306f\u6982\u89b3\u304c\u5927\u304d\u304f\u7570\u306a\u308b\u304c\u3001\u76f8\u540c\u306a\u5668\u5b98\u3067\u3042\u308b\u3002<\/p>\n\n\n\n

\u30a4\u30cc\u30b5\u30d5\u30e9\u30f3\u3067\u306f\u3001\u524d\u5e74\u30b7\u30e5\u30fc\u30c8\u306e\u830e\u8ef8\u304c\u80a5\u5927\u3057\u3066\u7403\u830e\u3068\u306a\u308b\u3002\u7403\u830e\u3092\u6398\u308a\u4e0a\u3052\u308b\u3068\u3001\u7b2c1\u8449\u306e\u8449\u9798\u306b\u7531\u6765\u3059\u308b\u5916\u5074\u306e\u76ae\u3068\u3001\u7b2c2\u8449\u306e\u8449\u9798\u306b\u7531\u6765\u3059\u308b\u5185\u5074\u306e\u76ae\u304c\u3042\u308b\u3002\u5f8c\u8005\u306f\u3001\u524d\u8005\u3088\u308a\u3082\u3084\u3084\u8272\u304c\u8584\u3044\u3053\u3068\u304c\u591a\u3044\u3002\u590f\u306e\u7d42\u308f\u308a\u306b\u3053\u308c\u3089\u306e\u76ae\u3092\u53d6\u308a\u9664\u304f\u3068\u3001\u7b2c1\u8449\u306e\u814b\u82bd\u3067\u3042\u308b\u66f4\u65b0\u82bd\u3068\u3001\u7b2c2\u8449\u306e\u814b\u82bd\u3067\u3042\u308b\u4e88\u5099\u82bd\u304c\u3059\u3067\u306b\u4f38\u9577\u3057\u3066\u3044\u308b\u3053\u3068\u304c\u308f\u304b\u308b\u3002<\/p>\n\n\n\n

\u30b0\u30ed\u30ea\u30aa\u30b5\u3067\u3082\u3001\u30a4\u30cc\u30b5\u30d5\u30e9\u30f3\u3068\u540c\u69d8\u306b\u524d\u5e74\u30b7\u30e5\u30fc\u30c8\u306e\u830e\u8ef8\u304c\u80a5\u5927\u3059\u308b\u3002\u3057\u304b\u3057\u3001\u30b0\u30ed\u30ea\u30aa\u30b5\u3067\u306f\u3001\u30a4\u30cc\u30b5\u30d5\u30e9\u30f3\u3068\u306f\u7570\u306a\u308a\u3001\uff12\u3064\u306e\u814b\u82bd\u306e\u57fa\u90e8\u304c\u68d2\u72b6\u306b\u4f38\u9577\u3059\u308b\u3002\u814b\u82bd\u306f\u3053\u306e\u68d2\u72b6\u90e8\u306e\u5148\u7aef\u306b\u4f4d\u7f6e\u3059\u308b\u3002\u68d2\u72b6\u90e8\u306f\u3001\u814b\u82bd\u306e\u57fa\u90e8\u306b\u3042\u308b\u524d\u5e74\u30b7\u30e5\u30fc\u30c8\u306e\u830e\u8ef8\u304c\u4f38\u9577\u3057\u305f\u3082\u306e\u3067\u3042\u308b\u305f\u3081\u3001\u8868\u9762\u306b\u306f\u8449\u306e\u75d5\u8de1\u304c\u306a\u304f\u3001\u814b\u82bd\u306e\u3042\u308b\u5148\u7aef\u90e8\u3092\u9664\u3044\u3066\u6ed1\u3089\u304b\u3067\u3042\u308b\u3002\u7fcc\u6625\u306b\u306a\u308b\u3068\u3001\u7b2c1\u8449\u306e\u814b\u82bd\u3067\u3042\u308b\u66f4\u65b0\u82bd\u304c\u4f38\u9577\u3057\u3066\u65b0\u3057\u3044\u30b7\u30e5\u30fc\u30c8\u3092\u5f62\u6210\u3059\u308b\u3002\u901a\u5e38\u306f\u7b2c1\u8449\u304a\u3088\u3073\u7b2c2\u8449\u306e\u814b\u82bd\u57fa\u90e8\u3060\u3051\u304c\u4f38\u9577\u3059\u308b\u305f\u3081\u3001\u4e8c\u53c9\u306e\u7403\u830e\u3068\u306a\u308b\u3002\u307e\u308c\u306b\u3001\u5199\u771f\u306b\u793a\u3059\u3088\u3046\u306b\u3001\u7b2c1\u8449\u3001\u7b2c2\u8449\u306e\u814b\u82bd\u306b\u52a0\u3048\u3066\u3001\u7b2c3\u8449\u306e\u814b\u82bd\u306e\u57fa\u90e8\u3082\u4f38\u9577\u3057\u3001\u4e09\u53c9\u306e\u7403\u830e\u3068\u306a\u308b\u5834\u5408\u304c\u3042\u308b\u3002<\/p>\n\n\n\n

Plants of Colchicaceae form corms or rhizomes (Nordenstam 1998). The axillary buds of the first two leaves of the current-year shoot become the shoot apices of the following year\u2019s shoots (Nordenstam 1998). Because the current-year shoot itself is also derived from an axillary bud, the first and second leaves correspond to prophylls, and the axillary buds of these prophylls become the shoot apices of the following year\u2019s shoots. In contrast, the axillary buds of leaves other than the prophylls form aerial branches. The axillary bud of the first leaf becomes the innovation bud and elongates in the following year. The axillary bud of the second leaf becomes the reserve bud, which usually elongates when the innovation bud is damaged or dies. However, depending on environmental conditions and species, both the innovation bud and the reserve bud may elongate. Although the corms of Colchicum<\/em> and Gloriosa<\/em> differ greatly in overall appearance, they are homologous organs.<\/p>\n\n\n\n

In Colchicum<\/em>, the stem axis of the previous year\u2019s shoot becomes swollen to form a corm. When the corm is dug up, it has an outer tunic derived from the leaf sheath of the first leaf and an inner tunic derived from the leaf sheath of the second leaf. The latter is often slightly paler than the former. When these tunics are removed at the end of summer, it can be seen that the innovation bud, which is the axillary bud of the first leaf, and the reserve bud, which is the axillary bud of the second leaf, have already begun to elongate. <\/p>\n\n\n\n

In Gloriosa<\/em>, as in Colchicum<\/em>, the stem axis of the previous year\u2019s shoot becomes swollen. However, unlike in Colchicum<\/em>, the bases of the two axillary buds elongate into rod-like structures in Gloriosa<\/em>. The axillary buds are located at the tips of these rod-like structures. Because each rod-like structure is formed by elongation of the stem axis of the previous year\u2019s shoot at the base of an axillary bud, its surface bears no leaf traces and is smooth except for the apical region bearing the axillary bud. In the following spring, the innovation bud, which is the axillary bud of the first leaf, elongates to form a new shoot. Usually, only the bases of the axillary buds of the first and second leaves elongate, producing a bifurcate corm. Rarely, as shown in the photograph, the base of the axillary bud of the third leaf also elongates, in addition to those of the first and second leaves, producing a trifurcate corm.<\/p>\n\n\n\n

\u5f15\u7528\u6587\u732e References<\/strong><\/p>\n\n\n\n

POWO (2026). Plants of the World Online<\/em>. Facilitated by the Royal Botanic Gardens, Kew. Published on the Internet; https:\/\/powo.science.kew.org\/<\/a>. Retrieved 3 June 2026. Distribution data from Kew Backbone Distributions (WCVP). \u00a9 World Checklist of Vascular Plants, licensed under CC BY 3.0.<\/p>\n\n\n\n

Brantjes, N.B.M., and Bos, J.J. (1980). Hawkmoth behavior and flower adaptation reducing self pollination in two Liliiflorae. New Phytol. 84<\/em>, 139\u2013143. https:\/\/doi.org\/10.1111\/J.1469-8137.1980.TB00756.X;WGROUP:STRING:PUBLICATION.<\/p>\n\n\n\n

Corbera, J., Alvarez-Cros, C., and Stefanescu, C. (2018). Evidence of butterfly wing pollination in the martagon lily Lilium martagon L. Butll. Inst. Catalana Hist. Nat. 82<\/em>, 117\u2013120.<\/p>\n\n\n\n

Corbera, J., Alvarez-Cros, C., and Stefanescu, C. (2018). Evidence of butterfly wing pollination in the martagon lily Lilium martagon L. Butll. Inst. Catalana Hist. Nat. 82<\/em>, 117\u2013120.<\/p>\n\n\n\n

Daniels, R.J., Johnson, S.D., and Peter, C.I. (2020). Flower orientation in Gloriosa superba (Colchicaceae) promotes cross-pollination via butterfly wings. Ann. Bot. 125<\/em>, 1137\u20131149. https:\/\/doi.org\/10.1093\/AOB\/MCAA048.<\/p>\n\n\n\n

Nordenstam, B. (1998). Colchicaceae. In The Families and Genera of Vascular Plants. Vol. III. Monocotyledons Lilianae (except Orchidaceae)., K. Kubitzki, ed. (Springer), pp. 175\u2013185.<\/p>\n\n\n\n

Thi, N. P.A., Kim, J.S., and Kim, J.H. (2013). Molecular phylogenetic relationships and implications for the circumscription of Colchicaceae (Liliales). Botanical Journal of the Linnean Society 172<\/em>, 255\u2013269. https:\/\/doi.org\/10.1111\/BOJ.12037;WGROUP:STRING:PUBLICATION.<\/p>\n\n\n\n

Weiss, M.R. (1997). Innate colour preferences and flexible colour learning in the<\/p>\n\n\n\n

pipevine swallowtail. Animal Behaviour 53<\/em>, 1043\u20131052.<\/p>\n","protected":false},"excerpt":{"rendered":"

\u30b0\u30ed\u30ea\u30aa\u30b5\u5c5e Gloriosa \u30b0\u30ed\u30ea\u30aa\u30b5\u5c5e\u306f\u30a4\u30cc\u30b5\u30d5\u30e9\u30f3\u79d1\u306b\u5c5e\u3057\u300111\u7a2e\u304c\u77e5\u3089 … \u7d9a\u304d\u3092\u8aad\u3080 →<\/span><\/a><\/p>\n","protected":false},"author":1,"featured_media":0,"comment_status":"closed","ping_status":"closed","sticky":false,"template":"","format":"standard","meta":[],"categories":[1],"tags":[],"_links":{"self":[{"href":"http:\/\/www.nibb.ac.jp\/plantdic\/blog\/index.php?rest_route=\/wp\/v2\/posts\/1577"}],"collection":[{"href":"http:\/\/www.nibb.ac.jp\/plantdic\/blog\/index.php?rest_route=\/wp\/v2\/posts"}],"about":[{"href":"http:\/\/www.nibb.ac.jp\/plantdic\/blog\/index.php?rest_route=\/wp\/v2\/types\/post"}],"author":[{"embeddable":true,"href":"http:\/\/www.nibb.ac.jp\/plantdic\/blog\/index.php?rest_route=\/wp\/v2\/users\/1"}],"replies":[{"embeddable":true,"href":"http:\/\/www.nibb.ac.jp\/plantdic\/blog\/index.php?rest_route=%2Fwp%2Fv2%2Fcomments&post=1577"}],"version-history":[{"count":1,"href":"http:\/\/www.nibb.ac.jp\/plantdic\/blog\/index.php?rest_route=\/wp\/v2\/posts\/1577\/revisions"}],"predecessor-version":[{"id":1586,"href":"http:\/\/www.nibb.ac.jp\/plantdic\/blog\/index.php?rest_route=\/wp\/v2\/posts\/1577\/revisions\/1586"}],"wp:attachment":[{"href":"http:\/\/www.nibb.ac.jp\/plantdic\/blog\/index.php?rest_route=%2Fwp%2Fv2%2Fmedia&parent=1577"}],"wp:term":[{"taxonomy":"category","embeddable":true,"href":"http:\/\/www.nibb.ac.jp\/plantdic\/blog\/index.php?rest_route=%2Fwp%2Fv2%2Fcategories&post=1577"},{"taxonomy":"post_tag","embeddable":true,"href":"http:\/\/www.nibb.ac.jp\/plantdic\/blog\/index.php?rest_route=%2Fwp%2Fv2%2Ftags&post=1577"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}