{"id":1066,"date":"2020-06-15T10:52:53","date_gmt":"2020-06-15T01:52:53","guid":{"rendered":"http:\/\/www.nibb.ac.jp\/plantdic\/blog\/?p=1066"},"modified":"2020-06-15T10:56:15","modified_gmt":"2020-06-15T01:56:15","slug":"%e3%82%a4%e3%82%b7%e3%83%a2%e3%83%81%e3%82%bd%e3%82%a6%e3%80%80drosera-lunata","status":"publish","type":"post","link":"http:\/\/www.nibb.ac.jp\/plantdic\/blog\/?p=1066","title":{"rendered":"\u30a4\u30b7\u30e2\u30c1\u30bd\u30a6\u3000Drosera lunata"},"content":{"rendered":"

\"Drosera<\/p>\n

\u30a4\u30b7\u30e2\u30c1\u30bd\u30a6\u8907\u5408\u7a2e\u7fa4Drosera peltata<\/em>\u00a0complex\u306f\u65e5\u672c\u304b\u3089\u897f\u306b\u97d3\u56fd\u3001\u4e2d\u56fd\u3001\u30a4\u30f3\u30c9\u30b7\u30ca\u3001\u30b9\u30ea\u30e9\u30f3\u30ab\u3001\u30a4\u30f3\u30c9\u3001\u30d2\u30de\u30e9\u30e4\u3078\u3001\u5357\u306b\u306f\u30d5\u30a3\u30ea\u30d4\u30f3\u3001\u30a4\u30f3\u30c9\u30cd\u30b7\u30a2\u3001\u30aa\u30fc\u30b9\u30c8\u30e9\u30ea\u30a2\u3001\u30cb\u30e5\u30fc\u30b8\u30fc\u30e9\u30f3\u30c9\u306b\u5206\u5e03\u3059\u308b\uff08Gibson et al. 2012)\u3002\u3057\u304b\u3057\u3001\u5206\u985e\u306f\u5341\u5206\u306b\u884c\u308f\u308c\u3066\u304a\u3089\u305a\u3001\u65e5\u672c\u3068\u30d2\u30de\u30e9\u30e4\u3067D. peltata<\/em>\u306e\u5b66\u540d\u3092\u3042\u3066\u3089\u308c\u3066\u3044\u308b\u30a4\u30b7\u30e2\u30c1\u30bd\u30a6\u306f\u3001\u30aa\u30fc\u30b9\u30c8\u30e9\u30ea\u30a2\u306eD. peltata<\/em>\u3068\u5f62\u614b\u304c\u7570\u306a\u308b\u306a\u3069\u306e\u6307\u6458\u304c\u3055\u308c\u3066\u304d\u305f\uff08Hara 1966)\u3002Gibson et al. (2012)\u306f\u5f62\u614b\u5f62\u8cea\u3092\u6bd4\u8f03\u3057\u3066\u3001\u30aa\u30fc\u30b9\u30c8\u30e9\u30ea\u30a2\u306e\u30a4\u30b7\u30e2\u30c1\u30bd\u30a6\u8907\u5408\u7a2e\u7fa4\u3092\u6574\u7406\u3059\u308b\u3068\u3068\u3082\u306b\u3001\u5f93\u6765D. peltata<\/em>\u3068\u8003\u3048\u3089\u308c\u3066\u304d\u305f\u30a4\u30f3\u30c9\u30cd\u30b7\u30a2\u304b\u3089\u6771\u30a2\u30b8\u30a2\u3078\u3068\u5206\u5e03\u3059\u308b\u7a2e\u306f\u3001D. lunata<\/em>\u3067\u3042\u308b\u3068\u767a\u8868\u3057\u305f\u3002\u65e5\u672c\u306e\u30a4\u30b7\u30e2\u30c1\u30bd\u30a6\u306b\u306f\u3001\u5f93\u6765Drosera peltata<\/em>\u306e\u5b66\u540d\u304c\u3042\u3066\u3089\u308c\u308b\u3053\u3068\u304c\u591a\u304b\u3063\u305f\u304c\u3001Drosera lunata<\/em>\u304c\u59a5\u5f53\u3060\u308d\u3046\u3068\u3044\u3046\u8ad6\u6587\u3067\u3042\u308b\u3002<\/p>\n

The Drosera peltate<\/em> complex is distributed from Japan, Korea, China, Indochine, Sri Lanka, India, the Himalayas, the Philippines, Indonesia, Australia, and New Zealand (Gibson et al. 2012). Gibson et al. (2012) compared morphological traits among the Drosera peltate<\/em> complex and classified into five species. The species distributed outside of Australia and New Zealand, which had been recognized as Drosera peltate<\/em>, was identified as D. lunata<\/em>.<\/p>\n

\"\"<\/p>\n

\u3053\u308c\u306f\u3001Gibson et al. (2012)\u306e\u30a4\u30b7\u30e2\u30c1\u30bd\u30a6\u8907\u5408\u7a2e\u7fa4\u306e\u691c\u7d22\u8868\u3067\u3042\u308b\u3002D. peltata<\/em>\u306f\u30ac\u30af\u7247\u306b\u6bdb\u304c\u3042\u308b\u304c\u3001\u65e5\u672c\u306e\u30a4\u30b7\u30e2\u30c1\u30bd\u30a6\u306b\u306f\u7121\u3044\u3002\u5f93\u3063\u3066\u3001D. peltata<\/em>\u3067\u306f\u306a\u3044\u53ef\u80fd\u6027\u304c\u9ad8\u3044\u3002\u4e00\u65b9\u3001\u30aa\u30fc\u30b9\u30c8\u30e9\u30ea\u30a2\u306eD. lunata<\/em>\u306f\u843c\u7247\u304c\u7121\u6bdb\u3067\u3042\u308b\u70b9\u304c\u3001\u65e5\u672c\u306e\u30a4\u30b7\u30e2\u30c1\u30bd\u30a6\u3068\u540c\u3058\u3067\u3042\u308b\u3002<\/p>\n

This is a key of the D. peltata<\/em> complex from Gibson et al. 2012. D. peltata<\/em> is defined to have hairs on sepals but Japanese species does not. Therefore, the Japanese species is not likely to be D. peltata<\/em>. On the other hand, D. lunata<\/em> from Australia has hairless sepals, which are the same as those in Japanese species.<\/p>\n

\"Drosera<\/p>\n

\u611b\u77e5\u770c\u6625\u65e5\u4e95\u5e02\u306b\u81ea\u751f\u3059\u308b\u30a4\u30b7\u30e2\u30c1\u30bd\u30a6\u306e\u82b1\u3092\u89b3\u5bdf\u3059\u308b\u3068\u3001\u843c\u7247\u80cc\u5074\uff08A\u3001\uff22\uff09\u306b\u306f\u591a\u6570\u306e\u817a\u7d30\u80de(C\u3001D\uff09\u304c\u3042\u308b\u304c\u3001\u6bdb\u306f\u751f\u3048\u3066\u3044\u306a\u3044\u3002\u843c\u7247\u5411\u8ef8\u5074\uff08\uff25\u3001\uff26\uff09\u306f\u6bdb\u3082\u817a\u7d30\u80de\u3082\u7121\u304f\u3001\u5e73\u6ed1\u3067\u3042\u308b\u3002Gibson et al. (2012)\u306f\u3001D. lunata<\/em>\u306e\u843c\u7247\u5f62\u614b\u3068\u3057\u3066\u3001\u300c\u843c\u7247\u306f\u5168\u7e01\u304b\u3089\u7d30\u92f8\u6b6f\u7e01\u3001\u3042\u308b\u3044\u306f\u3001\u7d30\u92f8\u6b6f\u72b6\u304b\u3089\u90e8\u5206\u7684\u306b0.6 mm\u307e\u3067\u306e\u9577\u7e01\u6bdb\u304c\u751f\u3048\u308b\u300d\u3068\u8a18\u8f09\u3057\u3066\u3044\u308b\u3002\u89b3\u5bdf\u3057\u305f\u30a4\u30b7\u30e2\u30c1\u30bd\u30a6\u3067\u306f\u3001\u8fba\u7e01\u306f\u7d30\u92f8\u6b6f\u72b6\u3068\u306a\u308a\u3001\u3055\u307e\u3056\u307e\u306a\u9577\u3055\u306e\u9577\u7e01\u6bdb\u304c\u5f62\u6210\u3055\u308c\u308b\u3002\u9577\u7e01\u6bdb\u306f\u3001\u9577\u3044\u3082\u306e\u3067\u7d040.4 mm\u3042\u308b\u3002\u5f93\u3063\u3066\u3001\u3053\u308c\u3089\u306e\u5f62\u8cea\u306f\u3001Gibson et al. (2012)\u306e\u8a18\u8f09\u306e\u7bc4\u56f2\u306b\u5165\u3063\u3066\u3044\u308b\u3002<\/p>\n

Sepals of D. lunata<\/em> collected in Kasugai City, Aichi Prefecture, Japan are glabrous with many unicellular\u00a0 glands on the abaxial side (A-D) but not on the adaxial side (E, F). Gibson et al. (2012) mentions “with the margin entire to serrulate, or denticulate, to partially to fully fimbriate with hairs up to 0.6mm long\u201d for the sepals of D. lunata<\/em>. The D. lunata<\/em> has characters in this range.<\/p>\n

\"Drosera<\/p>\n

\u6625\u65e5\u4e95\u5e02\u306e\u30a4\u30b7\u30e2\u30c1\u30bd\u30a6\u306e\u584a\u830e\u8868\u9762\u306f\u3001\u767d\u8272\u3067\u3042\u308b\u3002Gibson et al. (2012)\u306eD. lunata<\/em>\u306e\u8a18\u8f09\u306b\u306f\u8d64\u304b\u3089\u767d\u3068\u8a18\u8f09\u304c\u3042\u308a\u3001\u305d\u306e\u7bc4\u7587\u306b\u5165\u3063\u3066\u3044\u308b\u3002\u65e5\u672c\u306e\u30a4\u30b7\u30e2\u30c1\u30bd\u30a6\u306f\u3001\u3053\u3053\u307e\u3067\u898b\u3066\u304d\u305f\u843c\u7247\u5f62\u614b\u3068\u584a\u830e\u8868\u9762\u306e\u8272\u306b\u52a0\u3048\u3001\u8449\u5f62\u614b\u3001\u82b1\u5f62\u614b\u306a\u3069\u3082Gibson et al. (2012)\u306eD. lunata<\/em>\u306e\u7bc4\u7587\u306b\u5165\u3063\u3066\u304a\u308a\u3001\u73fe\u72b6\u3067\u306f\u3001D. lunata<\/em>\u3068\u3044\u3046\u5b66\u540d\u304c\u59a5\u5f53\u3067\u3042\u308b\u3068\u8003\u3048\u3089\u308c\u308b\u3002<\/p>\n

\u4e00\u65b9\u3067\u3001D. lunata<\/em>\u306f\u5e83\u57df\u5206\u5e03\u3059\u308b\u3053\u3068\u304b\u3089\u3001\u5730\u57df\u9593\u3067\u907a\u4f1d\u7684\u4ea4\u6d41\u304c\u906e\u3089\u308c\u3001\u5730\u57df\u96c6\u56e3\u3084\u96a0\u853d\u7a2e\u304c\u5f62\u6210\u3055\u308c\u3066\u3044\u308b\u53ef\u80fd\u6027\u304c\u3042\u308b\u3002\u843c\u7247\u5f62\u614b\u3068\u584a\u830e\u8868\u9762\u306e\u8272\u3067\u898b\u305f\u3088\u3046\u306b\u3001D. lunata<\/em>\u306e\u5f62\u614b\u5909\u7570\u306e\u7bc4\u56f2\u306f\u5927\u304d\u3044\uff08Gibson et al. 2012)\u3002\u4e00\u65b9\u3067\u3001\u65e5\u672c\u306e\u30a4\u30b7\u30e2\u30c1\u30bd\u30a6\u306e\u843c\u7247\u5f62\u614b\u3092\u4e2d\u56fd\u306e\u500b\u4f53(Lu and Kondo 2001)\u306e\u843c\u7247\u5f62\u614b\u3068\u6bd4\u8f03\u3059\u308b\u3068\u4e92\u3044\u306b\u985e\u4f3c\u3057\u3066\u3044\u308b\u304c\u3001\u30aa\u30fc\u30b9\u30c8\u30e9\u30ea\u30a2\u306e\u500b\u4f53\uff08Lowrie 2013)\u306f\u843c\u7247\u304c\u307b\u307c\u5168\u7e01\u3067\u5148\u7aef\u90e8\u5206\u306b\u7d30\u92f8\u6b6f\u3092\u6301\u3064\u70b9\u3067\u7570\u306a\u3063\u3066\u3044\u308b\u3002Lu and Kondo (2001)\u306b\u584a\u830e\u8868\u9762\u306e\u8272\u306e\u8a18\u8f09\u306f\u7121\u3044\u304c\u3001\u30aa\u30fc\u30b9\u30c8\u30e9\u30ea\u30a2\u306e\u500b\u4f53\u306f\u8d64\u8272\u3067\u3042\u308a\u3001\u65e5\u672c\u306e\u500b\u4f53\u304c\u767d\u8272\u3067\u3042\u308b\u306e\u3068\u7570\u306a\u3063\u3066\u3044\u308b\uff08Gibson et al. 2012\u3001Lowrie 2013)\u3002<\/p>\n

The surface of the tuber of the D. lunata<\/em>\u00a0in Kasugai is white. That in\u00a0D . lunata<\/em> sensu Gibson et al. (2012) is described to be red to white, and the surface color of Japanese D. lunata<\/em>\u00a0falls into the described characters. As well as the sepal morphology and tuber surface color, leaf and flower morphology is not distinguished Japanese D. lunata<\/em>\u00a0and D. lunata<\/em> sensu Gibson et al. (2012). Therefore, at present, D. lunata<\/em> should be an appropriate scientific name for the Japanease species.<\/p>\n

On the other hand, the widespread distribution of D. lunata<\/em> may interrupt genetic exchanges between regions, which can lead to regional populations and cryptic species. The range of morphological variation in D. lunata<\/em> is large, as seen in calyx morphology and tuber surface color (Gibson et al. 2012). While the sepal morphology of Chinese D. lunata<\/em>\u00a0(Lu and Kondo 2001) is similar to Japanese one, they are different from Australian plants (Lowrie 2013), having entire leaf margin with a slightly\u00a0serrated tip. The surface color of tuber in Australian D. lunata<\/em> is in red, which is diffenret from that in Japanese one (Gibson et al. 2012, Lowrie 2013). The surface color in Chinese species is not describe in Lu and Kondo 2001.<\/p>\n

\"Drosera<\/p>\n

\u30a4\u30b7\u30e2\u30c1\u30bd\u30a6\u8907\u5408\u7a2e\u7fa4\u542b\u3080\u3001\u584a\u830e\u3092\u6301\u3064\u30e2\u30a6\u30bb\u30f3\u30b4\u30b1\u306e\u4ef2\u9593\u3067\u3001\u8d64\u9053\u3092\u8d8a\u3048\u3066\u5317\u534a\u7403\u306b\u307e\u3067\u5206\u5e03\u3092\u5e83\u3052\u3066\u3044\u308b\u306e\u306fD. lunata<\/em>\u3060\u3051\u3067\u3042\u308b\u3002\u30aa\u30fc\u30b9\u30c8\u30e9\u30ea\u30a2\u3067\u306f\u3001\u7acb\u3061\u4e0a\u304c\u308b\u30b7\u30e5\u30fc\u30c8\u3092\u5f62\u6210\u3059\u308b\u584a\u830e\u6027\u30e2\u30a6\u30bb\u30f3\u30b4\u30b1\u306f\u3001\u65e9\u6625\u306b\u767a\u82bd\u3057\u3001\u6625\u304b\u3089\u590f\uff088\u304b\u308911\u6708\u9803\uff09\u306b\u958b\u82b1\u3059\u308b\u3053\u3068\u304b\u3089\uff08Lowrie 2013)\u3001\u958b\u82b1\u306f\u9577\u65e5\u3067\u8a98\u5c0e\u3055\u308c\u308b\u3068\u63a8\u6e2c\u3055\u308c\u308b\u3002\u4e00\u65b9\u3001D. lunata<\/em>\u306f\u3001\u590f\u671f\u306b\u6e7f\u3063\u305f\u5834\u6240\u3067\u751f\u80b2\u3057\u3001\u79cb\u304b\u3089\u6625\u306b\u304b\u3051\u3066\u958b\u82b1\u3059\u308b\u3053\u3068\u304c\u591a\u304f\u3001\u4ed6\u7a2e\u3068\u7570\u306a\u3063\u3066\u3044\u308b\uff08Gibson et al. 2012)\u3002\u3053\u306e\u3053\u3068\u304b\u3089\u3001\u9577\u65e5\u6027\u3092\u5931\u3063\u3066\u3044\u308b\u53ef\u80fd\u6027\u304c\u3042\u308b\u3002\u3055\u3089\u306b\u3001Gibson et al. (2012\uff09\u306f\u3001\u4e16\u754c\u5404\u5730\u306e\u6a19\u672c\u3092\u8abf\u3079\u308b\u3068\u3001\u958b\u82b1\u6642\u671f\u304c\u3055\u307e\u3056\u307e\u3067\u3042\u308b\u3068\u8a18\u8ff0\u3057\u3066\u304a\u308a\u3001\u9577\u65e5\u6027\u3092\u5931\u3063\u3066\u3044\u308b\u53ef\u80fd\u6027\u3092\u652f\u6301\u3057\u3066\u3044\u308b\u3002\u9577\u65e5\u6027\u3092\u5931\u3048\u3070\u3001\u65e5\u9577\u5909\u5316\u306e\u7121\u3044\u71b1\u5e2f\u57df\u3067\u306e\u958b\u82b1\u3092\u4ecb\u3057\u305f\u6709\u6027\u751f\u6b96\u304c\u53ef\u80fd\u3068\u306a\u308b\u3002\u3053\u308c\u3089\u306e\u3053\u3068\u304b\u3089\u3001D. lunata<\/em>\u306f\u82b1\u82bd\u5f62\u6210\u306e\u5236\u5fa1\u6a5f\u69cb\u306b\u5909\u7570\u304c\u304a\u3053\u308b\u3053\u3068\u3067\u3001\u71b1\u5e2f\u57df\u3068\u3044\u3046\u4e2d\u65e5\u6761\u4ef6\u3067\u306e\u7e41\u6b96\u3092\u53ef\u80fd\u3068\u3057\u3066\u3001\u5317\u534a\u7403\u3078\u3068\u5206\u5e03\u3092\u5e83\u3052\u308b\u3053\u3068\u3092\u53ef\u80fd\u306b\u3057\u305f\u306e\u3067\u306f\u306a\u3044\u304b\u3068\u3044\u3046\u4eee\u8aac\u3092\u305f\u3066\u308b\u3053\u3068\u304c\u3067\u304d\u308b\u3002\u5357\u5317\u534a\u7403\u3068\u71b1\u5e2f\u57df\u306b\u5206\u5e03\u3059\u308bD. lunata<\/em>\u3092\u3001\u7570\u306a\u3063\u305f\u65e5\u9577\u4e0b\u3067\u6bd4\u8f03\u683d\u57f9\u3057\u3001\u65e5\u9577\u3068\u82b1\u82bd\u8a98\u5c0e\u306e\u95a2\u4fc2\u3092\u8abf\u3079\u308c\u3070\u3053\u306e\u4eee\u8aac\u3092\u691c\u8a3c\u3067\u304d\u308b\u3060\u308d\u3046\u3002\u690d\u7269\u306f\u65e5\u9577\u3092\u5229\u7528\u3057\u3066\u958b\u82b1\u6642\u671f\u3084\u4f11\u7720\u6642\u671f\u3092\u6700\u9069\u5316\u3057\u3066\u3044\u308b\u3068\u3068\u3082\u306b\u3001\u305d\u308c\u304c\u7a2e\u5f62\u6210\u306e\u539f\u52d5\u529b\u306b\u3082\u306a\u3063\u3066\u3044\u308b\uff08Serrano-Bueno et al. 2017\u3001Gaudinier and Blackman 2020)\u3002\u4e00\u65b9\u3067\u3001\u3053\u306e\u3053\u3068\u306f\u3001\u5206\u5e03\u3092\u5e83\u3052\u308b\u5236\u7d04\u3068\u306a\u3063\u3066\u3044\u308b\u3002\u3053\u308c\u3089\u306e\u8981\u56e0\u306e\u517c\u306d\u5408\u3044\u306b\u3088\u3063\u3066\u3001\u591a\u69d8\u306a\u690d\u7269\u306e\u5206\u5e03\u304c\u751f\u307f\u51fa\u3055\u308c\u3066\u3044\u308b\u3068\u8003\u3048\u3089\u308c\u308b\u3002<\/p>\n

In tuberous Drosera<\/em>\u00a0species including D. peltata<\/em> comples, only D. lunata<\/em> expanded its distribution to the northern hemisphere across the equator. Tuber Drosera<\/em> species with erect leafy shoots start to grow from dormant tubers in early spring after the dormancy starting from dry summer. They bloom in spring to early summer (August to November), suggesting that flowering is induced by long days. D. lunata<\/em>, on the other hand, grows in wet areas during the summer months and often blooms in the fall and spring , different from other species (Gibson et al. 2012). This suggests that D. lunata may have lost its long-day nature. Furthermore, Gibson et al. (2012) examined specimens from around the world and found that the flowering season is variable, which supports the hypothesis. Loss of long-day nature would allow sexual reproduction via flowering in the tropics, where there is mostly no change in daylength. These results suggest that D. lunata<\/em> may be able to reproduce sexually via flowering in the equatorial region by mutations in flowering genes. Plants use day length to optimize flowering and dormancy times, which sometimes functions in speciation (Serrano-Bueno et al. 2017, Gaudinierand Blackman 2020). On the other hand, this is a constraint on expanding the distribution. The combination of these factors is likely to produce a diverse distribution patterns of plants.<\/p>\n

\"Drosera\u30a4\u30b7\u30e2\u30c1\u30bd\u30a6\u306e\u584a\u830e\uff08\u767d\u77e2\u5370\uff09\u306f\uff15\u301c\uff11\uff10\u30bb\u30f3\u30c1\u306e\u6df1\u3055\u306b\u3042\u308b\u3002\u830e\u304c\u4f38\u3073\u305f\u9803\u306b\u306f\u4eca\u5e74\u306e\u7403\u830e\u306f\u9ed2\u304f\u67af\u308c\u3066\u3044\u308b\u3002\u830e\u306e\u5730\u4e0b\u90e8\u306e\u9c57\u7247\u8449\u306e\u8449\u814b\u4e21\u5074\uff08\u8d64\u77e2\u5370\uff09\u3068\u9c57\u7247\u8449\u306e\u5916\u5074\uff08\u7d2b\u77e2\u5370\uff09\u304b\u3089\u4e0d\u5b9a\u6839\u304c\u51fa\u308b\u3002\u306a\u305c\u3053\u306e\u4f4d\u7f6e\u304b\u3089\u51fa\u308b\u304b\u306f\u308f\u304b\u3089\u306a\u3044\u3002<\/p>\n

The tuber of\u00a0D. lunata<\/em> is in 5 to 10 cm depth. By the time the stem has grown, this year\u2019s tuber turns black and mostly dead. The adventitious roots emerge from both sides of the leaf axils (red arrows) and from the adaxial side of the base of scale leaves (purple arrows).<\/p>\n

\"Drosera<\/p>\n

6\u6708\u521d\u65ec\u306b\u3001\u584a\u830e\u306e\u9ed2\u3044\u76ae\u3092\u3080\u304f\u3068\u53ce\u7e2e\u3057\u3066\u9ed2\u304f\u306a\u3063\u305f\u584a\u830e\uff08\u767d\u77e2\u5370\uff09\u3068\u767d\u3044\u584a\u830e\u304c\u3042\u3063\u305f\u3002<\/p>\n

In early June, the black skin of the tuber was peeled back to reveal a shrunken, blackened tuber (white arrow) and a white tuber.<\/p>\n

\"Drosera<\/p>\n

\u9ed2\u304f\u7e2e\u3093\u3060\u584a\u830e\u306f\u4eca\u5e74\u306e\u584a\u830e\u3067\u3001\u3053\u3053\u304b\u3089\u5730\u4e0a\u30b7\u30e5\u30fc\u30c8\u304c\u4f38\u9577\u3057\u3066\u3044\u308b\uff08\u767d\u77e2\u5370\u7dda\u3002\u77e2\u5370\u65b9\u5411\u304c\u5148\u7aef\uff09\u3002\u4e00\u65b9\u3001\u5730\u4e0a\u30b7\u30e5\u30fc\u30c8\u306e\u57fa\u90e8\u306e\u8449\uff08\uff0a\uff09\u306e\u814b\u304b\u3089\u9c57\u7247\u8449\u306e\u307f\u304b\u3089\u306a\u308b\u5074\u679d\u304c\u4f38\u9577\u3057\uff08\u8d64\u77e2\u5370\u7dda\uff09\u3001\u80a5\u5927\u3059\u308b\u3053\u3068\u3067\u3001\u767d\u3044\u584a\u830e\u3001\u3059\u306a\u308f\u3061\u3001\u6765\u5e74\u306e\u584a\u830e\u3092\u5f62\u6210\u3059\u308b\u3002<\/p>\n

The black, shrunken tuber is this year’s tuber, from which the above-ground shoot grows (white arrow line. The arrow direction is the tip). On the other hand, a lateral branch subtending only scale-leaves extends from the axils of a scale-leaf (*) at the base of the above-ground shoot (red arrow line) and it forms a white tuber, i.e., next year’s tuber.<\/p>\n

\"Drosera\u5074\u679d\u306e\u830e\u306f\u307b\u3068\u3093\u3069\u4f38\u9577\u305b\u305a\u3001\u5c0f\u3055\u306a\u9c57\u7247\u72b6\u306e\u8449\u304c\u5f62\u6210\u3055\u308c\u308b\uff08\u70b9\u7dda\uff09\u3002\u524d\u56f3\u306e\u3088\u3046\u306b\u3001\u6700\u521d\u3001\u5074\u679d\u306e\u830e\u9802\u306f\u4e0b\u5411\u304d\u306b\u51fa\u308b\u304c\u3001\u9014\u4e2d\u3067\u5411\u304d\u304c\u53cd\u8ee2\u3057\u3066\u3001\u5730\u4e0a\u5074\u3092\u5411\u304f\u3002\u830e\u9802\u90e8\u5206\uff08\u77e2\u5370\uff09\u306b\u306f\u591a\u6570\u306e\u9c57\u7247\u8449\u304c\u5f62\u6210\u3055\u308c\u308b\u3002\u767d\u661f\u5370\u306f\u4eca\u5e74\u306e\u584a\u830e\u306e\u4ed8\u3044\u3066\u3044\u305f\u4f4d\u7f6e\u3067\u3001\u5468\u8fba\u306b\u8d64\u307f\u304c\u304b\u3063\u305f\u9c57\u7247\u8449\u304c\u898b\u3048\u308b\u3002<\/p>\n

The stem of the lateral branch barely elongates, and small\u00a0scaly leaves are formed (dotted lines). As shown in the previous figure, the shoot apex of the lateral branch initially grows downward, but the direction reverses midway and faces toward the ground. Numerous scale leaves are formed at the stem apex (arrow). The white star indicates the position where this year’s tuber was attached, and reddish scale leaves can be seen in the vicinity.<\/p>\n

 <\/p>\n

\"Drosera\u584a\u830e\u3092\u7e26\u5272\u308a\u306b\u3059\u308b\u3068\u3001\u7dad\u7ba1\u675f(\u9ed2\u70b9\u7dda\uff09\u304c\uff35\u5b57\u578b\u306b\u306a\u308a\u3001\u830e\u9802\u304c\u9014\u4e2d\u304b\u3089\u4e0a\u3092\u5411\u3044\u3066\u3044\u308b\u306e\u304c\u308f\u304b\u308b\u3002<\/p>\n

When the tuber is longitudinally cut, the vascular bundle (black dotted line) is observed to be in U-shape, which indicates that the shoot apex changed the growth directions.<\/p>\n

\"DroseraDrosera peltata\u00a0sensu<\/em> Vickery 1933\u3067\u306f\u3001\u6765\u5e74\u306e\u584a\u830e\u4ee5\u5916\u306b\u3001\u5730\u4e0a\u30b7\u30e5\u30fc\u30c8\u306e\u5730\u4e0a\u90e8\u57fa\u90e8\u3084\u5730\u4e0b\u90e8\u306e\u9c57\u7247\u8449\u814b\u304b\u3089\u3001\u30c9\u30ed\u30c3\u30d1\u30fcdropper\u3068\u547c\u3070\u308c\u308b\u3001\u6570\u30bb\u30f3\u30c1\u307b\u3069\u4f38\u9577\u3059\u308b\u5074\u679d\u304c\u5f62\u6210\u3055\u308c\u308b\uff08Vickery 1933)\u3002\u30c9\u30ed\u30c3\u30d1\u30fc\u306f\u6700\u521d\u306f\u4e0b\u65b9\u306b\u5411\u304b\u3063\u3066\u4f38\u3073\uff08\u5de6\u56f3\u3001\u8d64\u77e2\u5370\u306f\u830e\u9802\u5206\u88c2\u7d44\u7e54\u3001\u9ed2\u7dda\u306f\u7dad\u7ba1\u675f\u3092\u793a\u3059\uff09\u3001\u5730\u4e2d\u3092\u3057\u3070\u3089\u304f\u4f38\u9577\u3057\u305f\u5f8c\u3001\u830e\u9802\u304c\u4e0a\u65b9\u306b\u5411\u304b\u3063\u3066\u66f2\u304c\u308b\u3053\u3068\u3067\u5074\u679d\uff08\u30c9\u30ed\u30c3\u30d1\u30fc\uff09\u5148\u7aef\u306b\u584a\u830e\u304c\u3067\u304d\u308b(\u53f3\u56f3\uff09\u3002<\/p>\n

In D. peltata sensu <\/em>Vickery 1933, aerial lateral branches called droppers are formed from axils of cauline leaves (Vickery 1933). Droppers grow downwards (left drawing) and enter into the soil. After a period of elongation in the ground, the stem apex bend upward to form a tuber (right drawing).<\/p>\n

\"DroseraDrosera lunata<\/em>\u3067\u306f\u3001\u5730\u4e0a\u830e\u304b\u3089\u306f\u30c9\u30ed\u30c3\u30d1\u30fc\u306f\u5f62\u6210\u3055\u308c\u306a\u3044\u304c\u3001\u5730\u4e0b\u306e\u9c57\u7247\u8449\u304b\u3089\u8907\u6570\u306e\u30c9\u30ed\u30c3\u30d1\u30fc\u304c\u5f62\u6210\u3055\u308c\u3001\u4eca\u5e74\u306e\u584a\u830e\u306e\u56de\u308a\u306b\u8907\u6570\u306e\u6765\u5e74\u306e\u584a\u830e\u304c\u5f62\u6210\u3055\u308c\u308b\u3002\u81ea\u751f\u5730\u3067\u306f\u3001\u4f55\u672c\u304b\u304c\u304b\u305f\u307e\u3063\u3066\u751f\u3048\u3066\u3044\u308b\u5834\u5408\u304c\u591a\u3044\u304c\u3001\u3053\u308c\u3089\u306f\u30c9\u30ed\u30c3\u30d1\u30fc\u306b\u3088\u308b\u6804\u990a\u7e41\u6b96\u3067\u5897\u3048\u305f\u3068\u8003\u3048\u3089\u308c\u308b\u3002<\/p>\n

Drosera lunata<\/em> does not form droppers on the aerial shoots but form from the axils of underground scale leaves. Since multiple droppers are formed to be next year’s tubers, several plants grow in clumps.<\/p>\n

\"Drosera\u30aa\u30fc\u30b9\u30c8\u30e9\u30ea\u30a2\u306b\u5206\u5e03\u3059\u308b\u584a\u830e\u3092\u6301\u3061\u5730\u4e0a\u830e\u304c\u4f38\u9577\u3059\u308b\u7a2e\u7fa4\u3067\u306f\u3001\u5730\u4e0a\u830e\u57fa\u90e8\u306e\u8449\u304c\u8449\u8eab\u3092\u5f62\u6210\u3059\u308b\u304b\u3001\u5f62\u6210\u305b\u305a\u306b\u9c57\u7247\u72b6\u306b\u306a\u308b\u304b\u304c\u3001\u7a2e\u3092\u533a\u5225\u3059\u308b\u91cd\u8981\u306a\u5f62\u8cea\u3068\u306a\u3063\u3066\u3044\u308b\u3002\u4e00\u65b9\u3001D. lunata<\/em>\u3067\u306f\u3001\u57fa\u90e8\u306b\u8449\u9593\u306e\u77ed\u3044\u30ed\u30bc\u30c3\u30c8\u8449\u3092\u6570\u679a\u5f62\u6210\u3059\u308b\u500b\u4f53\u3068\u3001\u8449\u8eab\u306e\u7121\u3044\u9c57\u7247\u72b6\u306e\u8449\u3092\u6570\u679a\u5f62\u6210\u3059\u308b\u500b\u4f53\u304c\u6df7\u5728\u3057\u3066\u3044\u308b\u3002\u3069\u3046\u3057\u3066\u3053\u306e\u3088\u3046\u306a\u9055\u3044\u304c\u751f\u3058\u308b\u306e\u304b\u306f\u308f\u304b\u3063\u3066\u3044\u306a\u3044\u3002<\/p>\n

In Australian tuber Drosera<\/em> with erect stems, leaf organization at the base of aerial shoots is a critical diagnostic character, although there are some variations: the existence of rosette leaves, the number of scale leaves, and so on. In D. lunata<\/em>, some plants form rosette leaves at the base of the aerial shoots, while others growing in proximate distances form scale leaves instead rosette leaves. The regulatory mechanisms to form different types of leaves are unknow in D. lunata<\/em>.\"Drosera<\/p>\n

\u5730\u4e0a\u30b7\u30e5\u30fc\u30c8\u306e\u57fa\u90e8\u306e\u30ed\u30bc\u30c3\u30c8\u8449\uff08\uff22\uff09\u3068\u5730\u4e0a\u30b7\u30e5\u30fc\u30c8\u306e\u305d\u308c\u4ee5\u5916\u306e\u8449\uff08\uff21\uff09\u3067\u306f\u5f62\u614b\u304c\u7570\u306a\u3063\u3066\u3044\u308b\u304c\u3001\u3053\u306e\u9055\u3044\u3092\u53f8\u308b\u6a5f\u69cb\u306f\u4e0d\u660e\u3067\u3042\u308b\u3002<\/p>\n

The morphology of the rosette leaf at the base of the aerial shoot (B) differs from that of the rest of the ground shoot (A), but the mechanism responsible for this difference is unknown.<\/p>\n

\"\u30cf\u30a8\u30c8\u30ea\u30bd\u30a6\u3000\u30b3\u30e2\u30a6\u30bb\u30f3\u30b4\u30b1\"<\/p>\n

\u8a2a\u82b1\u6606\u866b\u306f\u82b1\u7c89\u3092\u5a92\u4ecb\u3059\u308b\u3053\u3068\u3067\u3001\u591a\u304f\u306e\u88ab\u5b50\u690d\u7269\u306e\u6709\u6027\u751f\u6b96\u306b\u5bc4\u4e0e\u3057\u3066\u3044\u308b\u3002\u3057\u304b\u3057\u3001\u98df\u866b\u690d\u7269\u306f\u6606\u866b\u3092\u6355\u98df\u3059\u308b\u306e\u3067\u3001\u8a2a\u82b1\u6606\u866b\u3092\u6355\u7372\u3059\u308b\u3068\u82b1\u7c89\u5a92\u4ecb\u306b\u30c7\u30e1\u30ea\u30c3\u30c8\u306b\u306a\u308b\u3068\u3044\u3046\u30b8\u30ec\u30f3\u30de\u3001\u5373\u3061\u3001\u9001\u7c89\u8005-\u7372\u7269\u76f8\u53cdpollinator\u2013prey conflict\u3092\u62b1\u3048\u3066\u3044\u308b\u3002\u3053\u306e\u76f8\u53cd\u3092\u56de\u907f\u3059\u308b\u305f\u3081\u3001\u9577\u3044\u82b1\u830e\u3092\u5f62\u6210\u3057\u3001\u82b1\u3092\u6355\u866b\u8449\u304b\u3089\u96e2\u3059\u3053\u3068\u3067\u3001\u9001\u7c89\u6606\u866b\u3092\u6355\u7372\u3057\u3066\u3057\u307e\u308f\u306a\u3044\u3088\u3046\u306b\u3057\u3066\u3044\u308b\u306e\u3067\u306f\u306a\u3044\u304b\u3068\u8003\u3048\u3089\u308c\u3066\u304d\u305f (Ellison and Gotelli 2001, Ellison et al. 2003, Givnish 1989, Juniper et al. 1989)\u3002\u5b9f\u969b\u3001\u30cf\u30a8\u30c8\u30ea\u30bd\u30a6\u3084\u30e2\u30a6\u30bb\u30f3\u30b4\u30b1\u306e\u4ef2\u9593\u306e\u591a\u304f\u306f\u9577\u3044\u82b1\u5e8f\u3092\u5f62\u6210\u3059\u308b\u3002<\/p>\n

Flower-visiting insects contribute to the sexual reproduction of many angiosperms by mediating pollen. However, since carnivorous plants prey on insects, capturing visiting insects is a disadvantage to pollen vectors. The dilemma is called pollinator-prey conflict. It has been hypothesized that to avoid this conflict, long inflorescence\u00a0stalks are formed not to capture pollinating insects (Ellison and Gotelli 2001, Ellison et al. 2003, Givnish 1989, Juniper et al. 1989). In fact, many members of Drosera<\/em> and its sister genus\u00a0Dionaea<\/em>\u00a0form long inflorescences.<\/p>\n

\"Drosera<\/p>\n

\u305d\u3053\u3067\u3001Anderson \uff082010\uff09\u306f\u540c\u6240\u7684\u306b\u81ea\u751f\u3059\u308b\u3001\u9577\u3044\u82b1\u830e\u3092\u4f38\u9577\u3055\u305b\u30ed\u30bc\u30c3\u30c8\u72b6\u306e\u6355\u866b\u8449\u3068\u82b1\u306e\u8ddd\u96e2\u304c\u96e2\u308c\u3066\u3044\u308bDrosera pauciflora<\/em>\u3068\u3001\u6355\u866b\u8449\u3068\u82b1\u304c\u8fd1\u63a5\u3057\u3066\u5f62\u6210\u3055\u308c\u308bDrosera cisfiflora<\/em>\u3067\u3001\u9001\u7c89\u6606\u866b\u304c\u6355\u7372\u3055\u308c\u5272\u5408\u3092\u6bd4\u8f03\u3057\u305f\u3002\u3057\u304b\u3057\u3001\u4e21\u8005\u3068\u3082\u306b\u3001\u9001\u7c89\u6606\u866b\u306f\u307b\u3068\u3093\u3069\u6355\u98df\u3055\u308c\u3066\u3044\u306a\u304b\u3063\u305f\u3002\u3053\u306e\u3053\u3068\u304b\u3089\u3001\u9577\u3044\u82b1\u830e\u306f\u3001\u4ed6\u306e\u975e\u98df\u866b\u690d\u7269\u3068\u540c\u3058\u3088\u3046\u306b\u3001\u8a2a\u82b1\u6606\u866b\u3092\u7372\u5f97\u3057\u3001\u7a2e\u5b50\u3092\u3088\u308a\u9060\u304f\u306b\u98db\u3070\u3059\u305f\u3081\u306e\u9069\u5fdc\u3067\u3042\u308a\u3001\u9001\u7c89\u8005\u3092\u6355\u98df\u3057\u306a\u3044\u305f\u3081\u3067\u306f\u306a\u3044\u3068\u7d50\u8ad6\u3065\u3051\u305f\u3002<\/p>\n

Then, Anderson (2010) compared D. cistiflora<\/em> forming flowers close to trap leaves and D. pauciflora<\/em> with a long scape, both of which grow in close proximity. Pollinators were rarely captured in trap leaves in both species and Anderson (2010) concluded that the long scape is adaptive to attract pollinators and to disperse seeds in Drosera as in other non-carnivorous plants, but not to trap pollinators in their carnivorous leaves.<\/p>\n

\"Drosera<\/p>\n

\u30a4\u30b7\u30e2\u30c1\u30bd\u30a6\u8907\u5408\u7a2e\u7fa4\u3067\u306f\u3001\u82b1\u304c\u6355\u866b\u8449\u306b\u304b\u306a\u308a\u8fd1\u63a5\u3057\u3066\u3044\u308b\u3002El-Sayed et al. (2016)\u306fDrosera auriculata<\/em>\u306e\u82b1\u3068\u6355\u866b\u8449\u304b\u3089\u653e\u51fa\u3055\u308c\u308b\u6606\u866b\u8a98\u5f15\u7269\u8cea\u3092\u89e3\u6790\u3057\u3001\u82b1\u306e\u8a98\u5f15\u7269\u8cea\u306f\u8a2a\u82b1\u6606\u866b\u3092\u547c\u3073\u5bc4\u305b\u308b\u304c\u3001\u8449\u306e\u8a98\u5f15\u7269\u8cea\u306f\u547c\u3073\u5bc4\u305b\u306a\u3044\u3053\u3068\u3092\u767a\u898b\u3057\u305f\u3002\u3053\u306e\u3053\u3068\u304b\u3089\u3001\u30a4\u30b7\u30e2\u30c1\u30bd\u30a6\u3067\u3082\u82b1\u3068\u6355\u866b\u8449\u3067\u5225\u3005\u306e\u5316\u5b66\u7269\u8cea\u3092\u4f5c\u3063\u3066\u3001\u7570\u306a\u3063\u305f\u6606\u866b\u3092\u8a98\u5f15\u3057\u3066\u3044\u308b\u53ef\u80fd\u6027\u304c\u3042\u308b\u3002<\/p>\n

Flowers of D. peltata<\/em> complex species are formed in fairly close proximity to trap leaves. In D. auriculate<\/em>, El-Sayed et al. (2016) showed that the chemicals released from the flowers and the trap leaves are different and may attract different insects not to make pollinators be attracted to trap leaves.<\/p>\n

\"Drosera<\/p>\n

\u4e00\u65b9\u3001\u30a4\u30b7\u30e2\u30c1\u30bd\u30a6\u306f\u81ea\u5bb6\u53d7\u7c89\u3067\u7d50\u5b9f\u3067\u304d\u308b\u3002\u591a\u304f\u306e\u82b1\u306b\u30a2\u30b6\u30df\u30a6\u30de\u304c\u82b1\u7c89\u3092\u98df\u3079\u306b\u6765\u3066\u3044\u305f\u304c\u3001\u30a2\u30b6\u30df\u30a6\u30de\u304c\u79fb\u52d5\u3059\u308b\u3053\u3068\u3067\u3001\u81ea\u5bb6\u53d7\u7c89\u304c\u884c\u308f\u308c\u3066\u3044\u305f\u3002\u30a4\u30b7\u30e2\u30c1\u30bd\u30a6\u306e\u4ed6\u82b1\u53d7\u7c89\u3068\u81ea\u5bb6\u53d7\u7c89\u306e\u5272\u5408\u3001\u30a2\u30b6\u30df\u30a6\u30de\u4ee5\u5916\u306e\u9001\u7c89\u6606\u866b\u306e\u7a2e\u985e\u3001\u30a2\u30b6\u30df\u30a6\u30de\u3068\u4ed6\u306e\u9001\u7c89\u8005\u306e\u9001\u7c89\u3078\u306e\u5bc4\u4e0e\u306e\u5272\u5408\u306f\u308f\u304b\u3063\u3066\u3044\u306a\u3044\u3002<\/p>\n

\u8b1d\u8f9e\u3000\u30a4\u30b7\u30e2\u30c1\u30bd\u30a6\u306e\u89b3\u5bdf\u306f\u3001\u540d\u57ce\u5927\u5b66\u68ee\u7530\u88d5\u5c06\u535a\u58eb\u3001\u57fa\u790e\u751f\u7269\u5b66\u7814\u7a76\u6240\u661f\u91ce\u6566\u535a\u58eb\u306b\u6750\u6599\u3092\u63d0\u4f9b\u3057\u3066\u3044\u305f\u3060\u304d\u307e\u3057\u305f\u3002\u5fc3\u3088\u308a\u611f\u8b1d\u7533\u3057\u4e0a\u3052\u307e\u3059\u3002<\/p>\n

D. lunata<\/em> can be self-pollinated. Thysanoptera insects were often observed in flowers to eat pollen, which function in self-pollination.<\/p>\n

I deeply appreciate Drs. Yasumasa Morita in Meijo University and Atsushi Hoshino in National Institute for Basic Biology for providing D. lunata and stimulating my curiosity.<\/p>\n

\u5f15\u7528\u6587\u732e\u3000References<\/p>\n

Anderson, B. (2010). Did Drosera evolve long scapes to stop their pollinators from being eaten? Ann. Bot. 106, 653\u2013657.<\/p>\n

Ellison AM, Gotelli NJ. Evolutionary ecology of carnivorous plants.\u00a0TREE.\u00a02001;16:623\u2013629.<\/p>\n

Ellison AM, Gotelli NJ, Brewer JS, et al. The evolutionary ecology of carnivorous plants.\u00a0Advances in Ecological Research.\u00a02003;33:1\u201374.<\/p>\n

El-Sayed, A.M., Byers, J.A., and Suckling, D.M. (2016). Pollinator-prey conflicts in carnivorous plants: When flower and trap properties mean life or death. Sci. Rep. 6, 21065.<\/p>\n

Gaudinier, A., and Blackman, B.K. (2020). Evolutionary processes from the perspective of flowering time diversity. New Phytol. 225, 1883\u20131898.<\/p>\n

Givnish TJ. Ecology and evolution of carnivorous plants. In: Abrahamson WG, editor.\u00a0Plant\u2013animal interactions.\u00a0New York: McGraw-Hill Inc; 1989. pp. 243\u2013290.<\/p>\n

Gibson, R., Conn, B.J., and Bruhl, J.J. (2012). Morphological evaluation of the Drosera peltata complex (Droseraceae). Aust. Syst. Bot. 25, 49\u201380.<\/p>\n

Hara, H. (1966) The flora of the eastern Himalaya: results of the botanical expedition to eastern Himalaya organized by the University of Tokyo 1960 and 1963. Univ. Tokyo Press, Tokyo.<\/p>\n

Juniper, B.E., Robins, R.J., and Joel, D.M. (1989). The Carnivorous Plants (London, Uk: Academic Press).<\/p>\n

Lowrie, A. 2013. Carnivorous Plants of Australia Magnum Opus Vol. 1. Poole, Dorset, UK: Redfern Natural History Productions.<\/p>\n

Lu, L. and Kondo, K. 2001. Droseraceae. In Wu, Z.Y. and Raven, P.H. eds. Flora of China Vol. 8. pp. 199-201.<\/p>\n

Serrano-Bueno, G., Romero-Campero, F.J., Lucas-Reina, E., Romero, J.M., and Valverde, F. (2017). Evolution of photoperiod sensing in plants and algae. Curr. Opin. Plant Biol. 37, 10\u201317.<\/p>\n

Vickery, J.W. (1933). Vegetative reproduction in Drosera paltata and D. auriculata. Proc. Linn. Soc. New South Wales 58, 245\u2013250.<\/p>\n","protected":false},"excerpt":{"rendered":"

\u30a4\u30b7\u30e2\u30c1\u30bd\u30a6\u8907\u5408\u7a2e\u7fa4Drosera peltata\u00a0complex\u306f\u65e5\u672c\u304b\u3089\u897f\u306b … \u7d9a\u304d\u3092\u8aad\u3080 →<\/span><\/a><\/p>\n","protected":false},"author":1,"featured_media":0,"comment_status":"closed","ping_status":"closed","sticky":false,"template":"","format":"standard","meta":[],"categories":[178,81,157,139,115,182,177],"tags":[],"_links":{"self":[{"href":"http:\/\/www.nibb.ac.jp\/plantdic\/blog\/index.php?rest_route=\/wp\/v2\/posts\/1066"}],"collection":[{"href":"http:\/\/www.nibb.ac.jp\/plantdic\/blog\/index.php?rest_route=\/wp\/v2\/posts"}],"about":[{"href":"http:\/\/www.nibb.ac.jp\/plantdic\/blog\/index.php?rest_route=\/wp\/v2\/types\/post"}],"author":[{"embeddable":true,"href":"http:\/\/www.nibb.ac.jp\/plantdic\/blog\/index.php?rest_route=\/wp\/v2\/users\/1"}],"replies":[{"embeddable":true,"href":"http:\/\/www.nibb.ac.jp\/plantdic\/blog\/index.php?rest_route=%2Fwp%2Fv2%2Fcomments&post=1066"}],"version-history":[{"count":5,"href":"http:\/\/www.nibb.ac.jp\/plantdic\/blog\/index.php?rest_route=\/wp\/v2\/posts\/1066\/revisions"}],"predecessor-version":[{"id":1090,"href":"http:\/\/www.nibb.ac.jp\/plantdic\/blog\/index.php?rest_route=\/wp\/v2\/posts\/1066\/revisions\/1090"}],"wp:attachment":[{"href":"http:\/\/www.nibb.ac.jp\/plantdic\/blog\/index.php?rest_route=%2Fwp%2Fv2%2Fmedia&parent=1066"}],"wp:term":[{"taxonomy":"category","embeddable":true,"href":"http:\/\/www.nibb.ac.jp\/plantdic\/blog\/index.php?rest_route=%2Fwp%2Fv2%2Fcategories&post=1066"},{"taxonomy":"post_tag","embeddable":true,"href":"http:\/\/www.nibb.ac.jp\/plantdic\/blog\/index.php?rest_route=%2Fwp%2Fv2%2Ftags&post=1066"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}